Educational Record

1994 Ph.D. Biological Sciences, Stanford University. Dissertation: "Effects of above- and belowground resource distribution on the ecology of vesicular-arbuscular mycorrhizas."

1988 Tropical Ecology field course, Organization for Tropical Studies, Costa Rica.

1986 Biology of Tropical Plants summer field course, Fairchild Tropical Gardens, Harvard University.

1985 B.A. (Biology), with Distinction , Swarthmore College.